Complex Social Interaction
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February 1, 2003. It was the breeding season and Marion, the alpha female was in heat. Tristan had no interest in Marion, she was Seneca's mate, but Marion was good at getting Tristan in 'trouble' with the boss.
In this photo Seneca is standing tall and threatening Tristan who is averting his gaze. Marion is on the ground snowing a mixture of submission and annoyed protest as she has ended up in the middle of this altercation between two high-ranking males.
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Location : Wolf Park
032004 gray wolf wolves social interaction behavior dominance submission breeding season snow alpha male beta male alpha female fussing protesting agonistic pucker submissive grin averting gaze